John Ottway (Liam Neeson) works in Alaska hunting the wolves that threaten an oil drilling team. On his last day on the job, Ottway pens a letter to his wife Ana (Anne Openshaw) and plans to commit suicide. While holding his gun to his mouth, however, Ottway hears the howl of a wolf, which stops him.
Upon the completion of the job, the team and Ottway embark on a plane home in a blizzard. The plane cannot withstand the weather and it crashes in the middle of nowhere. Ottway awakens and finds Todd Flannery (Joe Anderson) along with Talget (Dermot Mulroney), Diaz (Frank Grillo), Hendrick (Dallas Roberts), Burke (Nonso Anozie), Hernandez (Ben Bray) and a dying Lewenden (James Badge Dale). Lewenden is mortally injured and Ottway knows he is going to die. He calms Lewenden, who eventually accepts his situation and dies peacefully.
Ottway assumes leadership of the group and sets the survivors to task collecting material for a fire. While Ottway is searching for wood, he sees a woman in need of help, but soon finds her being eaten by a gray wolf who also attacks Ottway. He is quickly rescued by the others and explains that they are most likely standing in the wolves' territory and are unwelcome. After getting a fire started, the survivors take turns keeping watch. While urinating during watch, Hernandez is killed by a wolf. The remaining survivors find his body in the morning and Ottway suggests they leave the crash site because they are wide open to attack. Diaz questions Ottway's leadership and begins defying his orders. Before they leave, Ottway and the others remove the wallets from many of the bodies with the intent of eventually returning them to surviving family members.
While hiking across the snow, Flannery falls behind and is killed by three wolves. The rest continue on and make camp in the woods, where tension between Ottway and Diaz comes to a head as the survivors create makeshift weaponry. Diaz threatens Ottway with his knife and elbows Talget in the face, but is ultimately disarmed and overtaken by Ottway. The wolves are close by and the survivors meet the alpha male wolf who sends an omega male to test Diaz. However, the survivors are able to kill the wolf and eat it. Diaz severs its head and throws it back to the wolves as a symbol of defiance. The group bonds over personal stories while Burke begins to suffer and hallucinate from the high elevation. A blizzard approaches the camp and Burke eventually falls unconscious and freezes to death.
Further in their travels, the dwindling survivors come across a high canyon wall within which, screened by trees, they identify a river and a possible route of escape. Ottway suggests having one person jump to the trees on the opposite side to secure a line as a means of traverse. Hendrick leaps to the trees and secures the finish end of the traverse. Diaz and Ottway make it across the line to join Hendrick, but Talget, who is afraid of heights, loses his glasses on the way and freezes out of fear. He reluctantly continues across, but soon finds himself caught in the makeshift rope by an accessorizing ring attached to his boot. Talget struggles to untangle himself, but the strained line breaks and he crashes through the trees to the ground. Badly wounded, Talget is attacked and dragged away by wolves. While attempting to save Talget, Diaz falls from the tree and severely injures his knee.
The three remaining continue their trek and make it to the river, but an exhausted Diaz can make it no further, preferring to stay and die than go on when he has nothing waiting for him at home. Hendrick tries to convince Diaz to carry on, but Ottway tells him it is futile. They give each other final goodbyes and Diaz is left on the riverside as crackling of branches caused by wolves are heard in the distance.
Ottway and Hendrick continue, but are soon chased by the pack again. Fleeing, they both fall in the river and Hendrick gets his foot stuck between underwater rocks and drowns despite Ottway's attempts to save him. Ottway then curses God and asks for his help, but receives no response. Ottway continues on, but seems to despair, dropping to his knees. He pulls out each wallet collected from the dead, examines the personal photographs inside them and places them in a pile against the snow, adding his own wallet and letter to the pile. While he is going through the wallets, the pack of wolves surround him, leading Ottway to discover that he has walked right into their den. He is soon spotted by the Alpha male and the other members of the pack back off. Ottway has a vision of his wife, wherein she is dying on a hospital bed.
In the final scene, Ottway decides to fight for his life, emptying the contents of the backpack he's carrying. He tapes miniature alcohol bottles to his fist, then smashes them and tapes a knife to his other fist, and recites his father's poem aloud. The film then ends with a Ottway & the Alpha lunging at each other, followed by a cut to black, and the film's credits. Along with the post-credits scene, the film's ending is open to interpretation and is the subject of much debate and controversy.
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